CuGenDBv2

Gene ID	Lsi01G017460
Organism	Lagenaria siceraria USVL1VR-Ls (Bottle gourd (USVL1VR-Ls) v1)
Description	DNA-directed RNA polymerase II subunit 4
Genome location	chr01:16266677..16274242
RNA-Seq Expression	Lsi01G017460
Synteny	Lsi01G017460
Gene Ontology terms	GO:0000288 - nuclear-transcribed mRNA catabolic process, deadenylation-dependent decay (biological process) GO:0006367 - transcription initiation from RNA polymerase II promoter (biological process) GO:0016567 - protein ubiquitination (biological process) GO:0031990 - mRNA export from nucleus in response to heat stress (biological process) GO:0034402 - recruitment of 3'-end processing factors to RNA polymerase II holoenzyme complex (biological process) GO:0045948 - positive regulation of translational initiation (biological process) GO:0000932 - P-body (cellular component) GO:0005665 - RNA polymerase II, core complex (cellular component) GO:0061630 - ubiquitin protein ligase activity (molecular function) GO:0031369 - translation initiation factor binding (molecular function) GO:0008270 - zinc ion binding (molecular function) GO:0003899 - DNA-directed 5'-3' RNA polymerase activity (molecular function) GO:0000166 - nucleotide binding (molecular function)
InterPro domains	IPR038324 - Rpb4/RPC9 superfamily IPR010997 - HRDC-like superfamily IPR006590 - RNA polymerase Rpb4/RPC9, core IPR005574 - RNA polymerase subunit RPB4/RPC9

Homology

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GenBank top hits

e value

%identity

Alignment

KAG6571896.1 E3 ubiquitin-protein ligase MPSR1, partial [Cucurbita argyrosperma subsp. sororia]

2.8e-64

65.84

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        ME++ISRY  +VVEV  DLPT  +IPKHPS IE IGLIFTLRRR +HF VSENGER+L++ +E  K+PLCD+QF AE+FN  D+SLFH LS SF+N+KP+
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN
        IDAE VADALIERWIEIRD+ NSRSSD+VF MTVNLTI ++VAR+EIG      GG+   +VPACG  + K L+RLE ++E + ICVICLDEL  KKK+N
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN

Query:  GR
        GR
Subjt:  GR

XP_011654386.1 RING finger protein 141 [Cucumis sativus]

9.8e-78

72.14

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        MEY+ISRYST  VEVDID+P   KIPKHPSPIELIGL+FTLRRRCN F V ENGERQLVE +EI KVPLCDHQFP E+FNG  +SLF YL  SFENY+ K
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR
        IDAE+VA  LIE+WIEIR+QVNSRS DKVFSMTVNLTI++VV ++ +G D + FGGE  R+VPAC + ++K LKRLEM+EEDER+CVICLDE+ KK+NGR
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR

Query:  V
        +
Subjt:  V

XP_016901804.1 PREDICTED: RING finger protein 141-like [Cucumis melo]

1.7e-77

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        MEY+IS+YST  VE+DIDLP S++I KHPSPIELIGL+FTLRRRCN F V ENGERQLVE +E  KVPLCDHQFP E+FNG DNSLF YL  SFENY+PK
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR
        IDAE+VA ALIE+WI IRDQ+NSRS  KVFSMTVNLTI++VV ++ IG D + FGGE FR VPAC + ++K LKRLE+++EDERIC+ICLDE++KK+NGR
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR

XP_022952293.1 ERAD-associated E3 ubiquitin-protein ligase HRD1-like [Cucurbita moschata]

1.4e-63

64.85

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        ME++ISRY  +VVEV  DLPT  +IPKHPS IE IGLIFTLRRR +HF VSENGER+L++ +E  K+PLCD+QF AE+FN  D+SLFH LS SF+N+KP+
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN
        IDAE VADALIERWIEIRD+ NSRSSD+VF MTVNLT+ ++VAR+EIG      GG+   +VPACG  + K L+RL+ ++E + ICVICLDEL  KKK+N
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN

Query:  GR
        GR
Subjt:  GR

XP_023511497.1 ERAD-associated E3 ubiquitin-protein ligase HRD1-like [Cucurbita pepo subsp. pepo]

4.7e-64

65.35

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        ME++ISRY ++VVEV  DLPT  +IPKHPS IE IGLIFTLRRR +HF VSENGER+L++ +E  K+PLCD+QF AE+FN  D+SLFH LS SF+N+KP+
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN
        IDAE VADALIERWIEIRD+ NSRSSD+VF MTVNLTI ++VAR+EIG      GG+   +VPACG  + K L+RL+ ++E + ICVICLDEL  KKK+N
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN

Query:  GR
        GR
Subjt:  GR

TrEMBL top hits

e value

%identity

Alignment

A0A0A0L236 RING-type domain-containing protein

4.7e-78

72.14

Show/hide

Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        MEY+ISRYST  VEVDID+P   KIPKHPSPIELIGL+FTLRRRCN F V ENGERQLVE +EI KVPLCDHQFP E+FNG  +SLF YL  SFENY+ K
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR
        IDAE+VA  LIE+WIEIR+QVNSRS DKVFSMTVNLTI++VV ++ +G D + FGGE  R+VPAC + ++K LKRLEM+EEDER+CVICLDE+ KK+NGR
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR

Query:  V
        +
Subjt:  V

A0A1S4E0Q9 RING finger protein 141-like

8.1e-78

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        MEY+IS+YST  VE+DIDLP S++I KHPSPIELIGL+FTLRRRCN F V ENGERQLVE +E  KVPLCDHQFP E+FNG DNSLF YL  SFENY+PK
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR
        IDAE+VA ALIE+WI IRDQ+NSRS  KVFSMTVNLTI++VV ++ IG D + FGGE FR VPAC + ++K LKRLE+++EDERIC+ICLDE++KK+NGR
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR

A0A5A7SNF1 RING finger protein 141-like

8.1e-78

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        MEY+IS+YST  VE+DIDLP S++I KHPSPIELIGL+FTLRRRCN F V ENGERQLVE +E  KVPLCDHQFP E+FNG DNSLF YL  SFENY+PK
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR
        IDAE+VA ALIE+WI IRDQ+NSRS  KVFSMTVNLTI++VV ++ IG D + FGGE FR VPAC + ++K LKRLE+++EDERIC+ICLDE++KK+NGR
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGR

A0A6J1GK53 ERAD-associated E3 ubiquitin-protein ligase HRD1-like

6.6e-64

64.85

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Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        ME++ISRY  +VVEV  DLPT  +IPKHPS IE IGLIFTLRRR +HF VSENGER+L++ +E  K+PLCD+QF AE+FN  D+SLFH LS SF+N+KP+
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN
        IDAE VADALIERWIEIRD+ NSRSSD+VF MTVNLT+ ++VAR+EIG      GG+   +VPACG  + K L+RL+ ++E + ICVICLDEL  KKK+N
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN

Query:  GR
        GR
Subjt:  GR

A0A6J1I8N9 ERAD-associated E3 ubiquitin-protein ligase HRD1-like

2.5e-63

64.36

Show/hide

Query:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK
        ME++ISRY  +VVEV  DLPT  +IPKHPS IE IGLIFTLR R +HF VSE+GER+L++ +E  K+PLCD+QF AE+FN  D+SLFH LS SF+N+KP+
Subjt:  MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPK

Query:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN
        IDAE VADALIERWIEIRD+ NSRSSD+VF MTVNLT+ ++VAR+EIG      GG+   +VPACG  + K L+RLE ++E + ICVICLDEL  KKK+N
Subjt:  IDAELVADALIERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDEL--KKKRN

Query:  GR
        GR
Subjt:  GR

SwissProt top hits

e value

%identity

Alignment

O15514 DNA-directed RNA polymerase II subunit RPB4

2.6e-09

39.58

Show/hide

Query:  VFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        VF K+L+Y  RFSR+KN + +  VR         +L    + +L +FEL  L NLCPET EE+ A++PSL  +GR  +DE ++++L+D+   + F+
Subjt:  VFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

O48890 DNA-directed RNA polymerase II subunit 4

2.8e-35

81.44

Show/hide

Query:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        +VFEKSL YVKRFSRYKNPDAVRQVRE        IL    R+QL EFELCVLGNLCPETVEEA+AMVPSLKTKGRAHDDEAIEKMLNDLSL+KRFE
Subjt:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

Q54S04 DNA-directed RNA polymerase II subunit rpb4

9.0e-10

36.46

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Query:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRF
        R F K+L Y ++FSRYKN  +++QVR               +  L EFE+  L NLCPE  +EA +++PSLK      +D+ ++ +L++LS +++F
Subjt:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRF

Q6DBA5 DNA-directed RNA polymerases IV and V subunit 4

6.1e-06

31.58

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Query:  FEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        ++++L YV+    Y NP +VR+V E               Y +++ E+CV+ N   E+V+E +A +PSLKTK +   ++ ++  L +LS +K+ E
Subjt:  FEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

Q9D7M8 DNA-directed RNA polymerase II subunit RPB4

2.6e-09

39.58

Show/hide

Query:  VFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        VF K+L+Y  RFSR+KN + +  VR         +L    + +L +FEL  L NLCPET EE+ A++PSL  +GR  +DE ++++L+D+   + F+
Subjt:  VFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

Arabidopsis top hits

e value

%identity

Alignment

AT4G15950.1 RNA polymerase II, Rpb4, core protein

4.3e-07

31.58

Show/hide

Query:  FEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        ++++L YV+    Y NP +VR+V E               Y +++ E+CV+ N   E+V+E +A +PSLKTK +   ++ ++  L +LS +K+ E
Subjt:  FEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

AT5G09920.1 RNA polymerase II, Rpb4, core protein

2.0e-36

81.44

Show/hide

Query:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE
        +VFEKSL YVKRFSRYKNPDAVRQVRE        IL    R+QL EFELCVLGNLCPETVEEA+AMVPSLKTKGRAHDDEAIEKMLNDLSL+KRFE
Subjt:  RVFEKSLHYVKRFSRYKNPDAVRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

Sequences

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CDS sequence

Show/hide CDS sequence

ATGGAATATCGAATCTCTCGATACTCAACTATGGTGGTAGAAGTAGATATAGATCTTCCAACGTCGAAAAAAATCCCAAAGCATCCATCACCAATCGAACTAATTGGCCT
GATCTTCACCCTCCGTCGTCGTTGCAACCACTTCCTAGTATCGGAGAATGGCGAACGACAATTAGTAGAGCATAAAGAAATACCAAAAGTTCCACTATGCGACCACCAAT
TCCCTGCTGAAATGTTCAATGGAGTGGACAATTCTCTATTCCATTACCTTTCTCATAGTTTCGAAAACTACAAACCCAAAATTGATGCAGAGCTAGTCGCTGATGCCCTA
ATCGAGCGGTGGATTGAAATAAGGGACCAGGTGAATTCAAGAAGCTCCGACAAAGTGTTTTCAATGACGGTAAATTTGACGATTTCGAATGTCGTGGCTCGACATGAAAT
TGGTAGTGACAGAAACGTATTCGGTGGAGAATATTTTAGGATCGTTCCAGCATGTGGCGAAGTAATTAGGAAAACGTTGAAGAGATTGGAGATGAAAGAAGAAGATGAAA
GAATTTGTGTGATATGTTTGGATGAATTGAAGAAAAAGAGGAATGGAAGAGTGTTTGAAAAGTCACTTCATTATGTAAAACGTTTTAGTCGTTATAAGAACCCTGATGCT
GTTAGACAAGTCCGAGAGTATCCTTATATAATAATATTTTTCATTTTATGGGATGGTGGTAGGTATCAGTTGGCAGAGTTTGAGCTCTGCGTACTTGGCAATCTTTGTCC
TGAAACAGTTGAGGAAGCTATTGCTATGGTACCGTCTTTAAAGACTAAAGGGAGAGCACATGACGACGAAGCTATTGAGAAGATGTTGAATGACCTTTCATTGATAAAAA
GATTTGAGTAG

mRNA sequence

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Protein sequence

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MEYRISRYSTMVVEVDIDLPTSKKIPKHPSPIELIGLIFTLRRRCNHFLVSENGERQLVEHKEIPKVPLCDHQFPAEMFNGVDNSLFHYLSHSFENYKPKIDAELVADAL
IERWIEIRDQVNSRSSDKVFSMTVNLTISNVVARHEIGSDRNVFGGEYFRIVPACGEVIRKTLKRLEMKEEDERICVICLDELKKKRNGRVFEKSLHYVKRFSRYKNPDA
VRQVREYPYIIIFFILWDGGRYQLAEFELCVLGNLCPETVEEAIAMVPSLKTKGRAHDDEAIEKMLNDLSLIKRFE

Lsi01G017460 (gene) of Bottle gourd (USVL1VR-Ls) v1 genome